Description Usage Arguments Details Value Author(s) References Examples
Sets up and executes a HiSSE model (Hidden State Speciation and Extinction) on a phylogeny and character distribution.
1 2 3 4 5 6  hisse(phy, data, f=c(1,1), turnover=c(1,2), eps=c(1,2),
hidden.states=FALSE, trans.rate=NULL, condition.on.survival=TRUE,
root.type="madfitz", root.p=NULL, includes.fossils=FALSE, k.samples=NULL,
strat.intervals=NULL, sann=TRUE, sann.its=1000, bounded.search=TRUE,
max.tol=.Machine$double.eps^.50, starting.vals=NULL, turnover.upper=10000, eps.upper=3,
trans.upper=100, restart.obj=NULL, ode.eps=0, dt.threads=1)

phy 
a phylogenetic tree, in 
data 
a matrix (or dataframe) with three columns. The first column containing the species names and the second and third containing the binary character information. Character "0" is on column 2 and chracter "1" is on column 3. A value of 0 means character absent and a value of 1 character present. The input of data follows a Pagel model. See 'Details'. 
f 
vector of length 2 with the estimated proportion of extant species in state 0 and 1 that are included in the phylogeny. A value of c(0.25, 0.5) means that 25 percent of species in state 0 and 50 percent of species in state 1 are included in the phylogeny. By default all species are assumed to be sampled. 
turnover 
a numeric vector indicating the number of free turnover parameters in the model. 
eps 
a numeric vector indicating the number of free extinction fraction parameters in the model. 
hidden.states 
a logical indicating whether the model includes a
hidden states. The default is 
trans.rate 
provides the transition rate model. See function

condition.on.survival 
a logical indicating whether the likelihood
should be conditioned on the survival of two lineages and the
speciation event subtending them (Nee et al. 1994). The default is 
root.type 
indicates whether root summarization follow the procedure described by FitzJohn et al. 2009, “madfitz” or HerreraAlsina et al. 2018, “herr_als”. 
root.p 
a vector indicating fixed root state probabilities. The
default is 
includes.fossils 
a logical indicating whether the tree contains fossil taxa. The default is 
k.samples 
a table of extinct individuals with sampled descendants. See details for how the table must be formatted. 
strat.intervals 
a table of extinct individuals with sampled descendants. See vignette for how the table must be formatted. 
sann 
a logical indicating whether a twostep optimization
procedure is to be used. The first includes a simulate annealing
approach, with the second involving a refinement using

sann.its 
a numeric indicating the number of times the simulated annealing algorithm should call the objective function. 
bounded.search 
a logical indicating whether or not bounds should
be enforced during optimization. The default is is 
max.tol 
supplies the relative optimization tolerance to

starting.vals 
a numeric vector of length 3 with starting values for the model for all areas and hidden states. Position [1] sets turnover, [2] sets extinction fraction, and [3] transition rates. 
turnover.upper 
sets the upper bound for the turnover parameters. 
eps.upper 
sets the upper bound for the eps parameters. 
trans.upper 
sets the upper bound for the transition rate parameters. 
restart.obj 
an object of class that contains everything to restart an optimization. 
ode.eps 
sets the tolerance for the integration at the end of a branch. Essentially if the sum of compD is less than this tolerance, then it assumes the results are unstable and discards them. The default is set to zero, but in testing a value of 1e8 can sometimes produce stable solutions for both easy and very difficult optimization problems. 
dt.threads 
sets the number of threads available to data.table. In practice this need not change from the default of 1 thread, as we have not seen any speedup from allowing more threads. 
This function sets up and executes a new and faster version of the HiSSE model. Note that the fourstate characterindependent model can be called from this command in addition to the twostate BiSSE model and the full characterdependent HiSSE model. See vignette on how to set this up.
The “trans.rate” input is the transition model and has an
entirely different setup than turnover rates and extinction fraction. See
TransMatMakerHiSSE
function for more details.
For the “root.type” option, we are currently maintaining the previous default of “madfitz”. However, it was recently pointed out by HerreraAlsina et al. (2018) that at the root, the individual likelihoods for each possible state should be conditioned prior to averaging the individual likelihoods across states. This can be set doing “herr_als”. It is unclear to us which is exactly correct, but it does seem that both “madfitz” and “herr_als” behave exactly as they should in the case of characterindependent diversification (i.e., reduces to likelihood of tree + likelihood of trait model). We've also tested the behavior and the likelihood differences are very subtle and the parameter estimates in simulation are nearly indistinguishable from the “madfitz” conditioning scheme. We provide both options and encourage users to try both and let us know conditions in which the result vary dramatically under the two root implementations. We suspect they do not.
For userspecified “root.p”, you should specify the probability for each state. If you are doing a hidden model, there will be four states: 0A, 1A, 0B, 1B. So if you wanted to say the root had to be state 0, you would specify “root.p = c(0.5, 0, 0.5, 0)”.
This code will completely replace the original hisse function in the next version.
hisse
returns an object of class hisse.fit
. This is a list with
elements:
$loglik 
the maximum negative loglikelihood. 
$AIC 
Akaike information criterion. 
$AICc 
Akaike information criterion corrected for samplesize. 
$solution 
a matrix containing the maximum likelihood estimates of the model parameters. 
$index.par 
an index matrix of the parameters being estimated. 
$f 
usersupplied sampling frequencies. 
$hidden.states 
a logical indicating whether hidden states were included in the model. 
$condition.on.surivival 
a logical indicating whether the likelihood was conditioned on the survival of two lineages and the speciation event subtending them. 
$root.type 
indicates the userspecified root prior assumption. 
$root.p 
indicates whether the userspecified fixed root probabilities. 
$phy 
usersupplied tree 
$data 
usersupplied dataset 
$trans.matrix 
the usersupplied transition matrix 
$max.tol 
relative optimization tolerance. 
$starting.vals 
The starting values for the optimization. 
$upper.bounds 
the vector of upper limits to the optimization search. 
$lower.bounds 
the vector of lower limits to the optimization search. 
$ode.eps 
The ode.eps value used for the estimation. 
Jeremy M. Beaulieu
Beaulieu, J.M, and B.C. O'Meara. 2016. Detecting hidden diversification shifts in models of traitdependent speciation and extinction. Syst. Biol. 65:583601.
FitzJohn R.G., Maddison W.P., and Otto S.P. 2009. Estimating traitdependent speciation and extinction rates from incompletely resolved phylogenies. Syst. Biol. 58:595611.
HerreraAlsina, L., P. van Els, and R.S. Etienne. 2018. Detecting the dependence of diversification on multiples traits from phylogenetic trees and trait data. Systematic Biology, 68:317328.
Maddison W.P., Midford P.E., and Otto S.P. 2007. Estimating a binary characters effect on speciation and extinction. Syst. Biol. 56:701710.
Nee S., May R.M., and Harvey P.H. 1994. The reconstructed evolutionary process. Philos. Trans. R. Soc. Lond. B Biol. Sci. 344:305311.
1 2 3 4 5 6 7 8 9 10 11 12 13 14 15  library(diversitree)
pars < c(0.1, 0.2, 0.03, 0.03, 0.01, 0.01)
set.seed(4)
phy < tree.bisse(pars, max.t=30, x0=0)
sim.dat < data.frame(names(phy$tip.state), phy$tip.state)
## Fit BiSSE equivalent:
trans.rates.bisse < TransMatMakerHiSSE(hidden.traits=0)
pp.bisse < hisse(phy, sim.dat, hidden.states=FALSE, turnover=c(1,2),
eps=c(1,2), trans.rate=trans.rates.bisse)
## Now fit HiSSE equivalent with a hidden state for state 1:
trans.rates.hisse < TransMatMakerHiSSE(hidden.traits=1)
pp.hisse < hisse(phy, sim.dat, hidden.states=TRUE, turnover=c(1,2,1,2),
eps=c(1,2,1,2), trans.rate=trans.rates.hisse)

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